This means that you are somewhere setting the content-length header attribute manually in your code which for security reasons will no longer work in 4.3.
Would be nice if “QA Guy” or his colleagues could update the Unity 4.3 release notes and the Unity docs with the as yet undocumented “security reasons” and breaking change that prevent existing code from working.
FYI: After digging around in the WWW code with reflection, I found this: