My animation will not play after I apply armature in Blender.

If i don’t click apply on my armature in Blender, then it warps my meshes in unity, but the animation still plays. Once I apply the armature the mesh is no longer warped, but the animations won’t play in the game or the preview window. It’s like they are stuck on the first frame. Any help is appreciated. Thank you!

I never applied the modifiers in blender. They get applied during export and everything was always working fine.