Two same fbx files works different

Hi there.

I faced a very strange problem. I copy a file A from Explorer to get a file B. The newfile B and the source file A behave differently. All inspector settings are the same but the .meta files are different! How can that happen?

file A: alt text
file B: alt text

the animation in file B cannot play.
file B has a warning in inspector:
alt text

More clues:
file B has transform:
alt text
file A:
alt text
As far as I know, their import Settings are exactly the same.

Sounds like there was some change in import settings for file B. I would try coping the meta file with the actual file, and see if that keeps them the same. Either that or try re importing File A. Just depends on what you want to accomplish.