URP: 2021 LTS: Objects that look great in Scene window turn to black when I hit "play"

See images below…I’ve read through so many bits of advice, most of which is out of date, and I’m at a complete loss here…looks great in editor, as soon as I go to play the scene…it’s the black ball of death…

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I assume it’s a custom shader? Or is it just geometry with a texture?

Maybe remove all lights except 1 directional and set it to realtime.

Maybe try a different quality tier if you still have multiple presets?

Thanks! Yes, custom shader. Unless I’ve missed something, there is only one directional light…but I will recheck that.