As the picture shows, the Transform shown on the Inspector suddenly becomes strange. It’s displayed in a wrong way, even telling me the w axis!
I don’t know if it has something to do with the problem, but when I was playing with my project and pressed Build Settings… → Switch Platform from Web Player to PC and Mac Standalone, I got a really strange Unity error. I think was something like Assembly… maybe this was the cause of the Transform problem, which sometimes shows the correct way, and sometimes this weird way.
You’re just in Debug mode. Normally this is accessed with the drop-down menu at the upper-right on an Inspector pane. However, your Inspector doesn’t say “Debug”, so this is probably a bug. See if you can get around it by switching to Debug and then Normal mode.